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Dr. David Maitland BSc PhD FRPS Nature, Science, Super-Macro, Videography A Zoology degree from St. Andrews University was followed by a PhD in Zoology from the University of New South Wales, Sydney, Australia. Photography featured prominently in my studies of animal form and function. I employed conventional 35mm macro photography along with transmission and scanning electron microscopy to discover how crabs breathe air through their legs and how maggots jump. I became a full time Nature photographer in 2006. I love to capture the quirkiness in a creature's character, its intricate details and the beauty and wonder of Nature.

“David’s background is that of an Academic and naturalist – but the use of Photography to elucidate the intricacies of nature from an academic view point has now been transformed into a new sense of enquiry - the pursuit of art and wonder in Nature. Winner of the European Wildlife Photographer of the Year in 2008, and numerous awards in the Veolia Wildlife Photographer of the Year - amongst many others - have established David’s position as one of the world’s foremost nature photographers. David’s quirky insight into nature’s little details, has led to his collaboration on Brian Cox’s 2013 “Wonders of Life” series on BBC2, as their specialist- and macro-photographer.” 2013: Appointed External Examiner for the University of Falmouth BA (Hons) Marine & Natural History Photography degree programme.

Commissions: A specialist in macro, super-macro and microscopy, employing state-of-the-art techniques, David is available for bespoke advertising commissions. Videography and still photography of the highest quality. e-mail: david@davidmaitland.com for full details. Fine Art Prints: Photographs by David Maitland are available as signed Fine Art prints. e-mail: david@davidmaitland.com for full details. Courses / Workshops: Specialist tailor-made “one-to-one” or “you-and-a-friend” courses. Or, why not join a small group for one, two, or three-day courses in Close-up and Macro photography and Microscopy? Lakenheath Fen and Thetford Forest are close by to serve as a lovely “practice ground”. e-mail: david@davidmaitland.com for full details. Students: David is available for tailored tuition to suit your needs, from portfolio reviews to one-to-one tuition. e-mail: david@davidmaitland.com for full details.

Number of awards: 1

Awards 2013 NIKON Small World Photomicrography Competition: “Mollusc Radula” receives Honourable Mention Award. www.nikonsmallworld.com Olympus Bioscapes “Three-wise-men” receives Honourable Mention Award. www.olympusbioscapes.com EUROPEAN Wildlife Photographer of the Year : - Two photographs “Starry Night” & “Spicules” both Highly Commended. www.gdtfoto.de

Awards 2012 VEOLA WILDLIFE PHOTOGRAPHER OF THE YEAR 2012 Runner-up in “Creative Visions” “Sands of Time” (Coral Sand). NIKON Small World Photomicrography Competition: Coral Sand wins 18th place. Mosquito gets an Image of Distinction award. www.nikonsmallworld.com EUROPEAN Wildlife Photographer of the Year 2012 : - Two photographs Highly Commended www.gdtfoto.de The ROYAL PHOTOGRAPHIC SOCIETY’S 155th International Print Exhibition: Two prints accepted for International exhibition.

Awards 2011 VEOLIA WILDLIFE Photographer of the Year: "Fading Beauty" is Specially Commended: In Praise of Plants NIKON Small World Photomicrography Competition: An Honorable Mention and 2 Images of Distinction BWPA British Wildlife Photography Awards:- Highly Commended - Living Landscape

Awards 2009 PDN PhotoAnnual 2009 Award Winner Bronze Medal Award: Royal Photographic Society 152nd International Print Exhibition: "Who Eats Who" EUROPEAN Wildlife Photographer of the Year 2009 : - Highly Commended www.gdtfoto.de IPA - International Photography Awards Special - Winner: 2nd place & "Best of Show" award IGPOY - International Garden Photographer of the year Portfolio Finalist: Plant Portraits

Number of pubs: 1

Commissions: A specialist in macro, super-macro and microscopy, employing state-of-the-art techniques, David is available for bespoke advertising commissions. Videography and still photography of the highest quality. e-mail: david@davidmaitland.com for full details. Fine Art Prints: Photographs by David Maitland are available as signed Fine Art prints (some limited, some open). Print media is the superb Hahnemuhle Photo Rag. Prints are supplied unframed only. e-mail: david@davidmaitland.com for full details.

Courses / Workshops: Nature & Macro: Specialist tailor-made “one-to-one” or “you-and-a-friend” nature photography courses. Or, why not join a small group for one, two, or three-day courses in Close-up and Macro photography? Lakenheath Fen and/or Thetford Forest are close-by to serve as a lovely “practice ground”. Super-macro: As a Multiple award winner in the Nikon Small World international photographic competition David can help you to improve your macro techniques. Learn how to do focus stacking, and take beautiful super-macro photographs of living things. e-mail: david@davidmaitland.com for full details.

Microscopy: David has a fully equipped microscopy "CSI" lab with Light and fluorescence microscopes capable of polarizing, phase-contrast, DIC, Dark field and fluorescence techniques, amongst others. e-mail: david@davidmaitland.com for full details. Students: David is available for tailored tuition to suit your needs, from portfolio reviews to one-to-one tuition. e-mail: david@davidmaitland.com for full details.

Number of Titles: 10

Reptiles

Bush-meat

Frogs

Design 1

Invertebrates

Imago

Light

Colour

Design 2

Micro/Super Macro

Number of Captions: 85

The Passion Flower Butterfly lays its eggs singly at the tip of the coiling tendrils of the Passion Vine, Passiflora pittieri, as a defense against ant predation. Rainforest, Braulio Carrillo National Park, Caribbean lowlands, northern Costa Rica

An endangered Gabon Black Colobus monkey, Colobus satanas anthacinus, awaiting preparation. Food Market, Libreville, Gabon.

BUSH-MEAT: In preparation for the bush meat trade, monkeys are thrown onto open fires to burn off their fur prior to being sold naked at food markets throughout central West Africa. Despite being banned, the consumption of bush meat remains popular and presents a serious threat to both wildlife and Humans: The wildlife is threatened with extinction and Humans run the risk of contracting deadly diseases like Ebola and HIV. With fur removed and remnants scraped away, a pile of naked monkeys and a porcupine (lower right) lie awaiting sale. Food Market, Libreville, Gabon.

In preparation for the bush-meat trade, monkeys are thrown onto open fires to burn off their fur prior to being sold naked at food markets throughout central West Africa. This is an endangered Gabon Black Colobus monkey, Colobus satanas anthacinus. Food Market, Libreville, Gabon.

Human skull mid-line suture - the convoluted joint between the bone plates which form the top of the skull

An impressive and dangerous snake, the Jumping Pit Viper, or Jumping Tommygoff - Atropoides nummifer - keeps its eye on me, while I keep my eye on it... Close-up of the vertical cat-like pupil of its eye. Chiquibul National Park, Belize, Central America.

Coiled tail tip of a False Chameleon lizard – Polychrus gutturosus - Rainforest, Braulio Carrillo National Park, Caribbean lowlands, Costa Rica

In preparation for the bush-meat trade, monkeys are thrown onto open fires to burn off their fur prior to being sold naked at food markets throughout central West Africa. Despite being banned, the consumption of bush-meat remains popular and presents a serious threat to both wildlife and Humans: The wildlife is threatened with extinction and Humans run the risk of contracting deadly diseases like Ebola and HIV. This is an endangered Gabon Black Colobus monkey, Colobus satanas anthacinus. Food Market, Libreville, Gabon.

Hard to swallow: A young common frog, Rana temporaria, attempts to swallow an incongruously large earthworm. The striped body of the Tiger or Brandling Earthworm, Eisenia foetida, looks alien to the frog. In this case the frog manages to swallow the worm only to have the worm crawl back out of the frog's stomach and mouth a few minutes later – just too big for its stomach. Garden pond, Calne, Wiltshire, UK.

Hug : - A pair of common frogs, Rana temporaria, in Amplexus - the male is behind clasping, the female, fat with eggs, tightly around her arm-pits. Calne, Wiltshire, UK.

A small Reed frog, Hyperolius sp. rests during the day inside the young unfurled whorl of an Arrow-root (Marantaceae) leaf. Loango National Park. Gabon.

A female Strawberry Poison-dart or Poison-arrow frog, Dendrobates pumilio, carries a single tadpole to a nursery pond in the base of a Bromeliad plant. This is a classic example of parental care in the animal kingdom. Braulio Carrillo National Park, Caribbean lowlands, northern Costa Rica.

The gills of a field mushroom light up as rays of light when back lit.

A translucent and bioluminescent planktonic polychaete worm, Tomopteris (Annelida, Polychaeta), snakes gracefully through the water. Tomopteris is one of the few marine animals able to make yellow luminescence. The luminescent material is made in the parapodia (the animal's paddle-shaped limbs). North Sea, UK .

An Eastern Snake-necked Turtle or Long-necked Tortoise, Chelodina longicollis, smiles as it breakes through the pond weed surface to take a breath of air after foraging along the swamp bed. Sydney, NSW, Australia

European Grass snake, Natrix natrix, coiled and alert. Garden wildlife. Calne, Wiltshire, UK

A gecko is silhouetted against a banana leaf. Sao Tome e Principe.

A Common frog, Rana temporaria, sits upon a fallen purple iris flower. Calne, Wiltshire, UK.

Lacewing, Chrysopa carnea, rests amongst young maple leaves. Calne, Wiltshire, UK

The exoskeletal skin of many scorpions takes on an unusual property as it hardens; it fluoresces brightly under ultraviolet light. The function of this fluorescence is unknown. This large tropical black species (probably the Asian Forest Scorpion, Heterometrus spinifer) is a good example. Tanjun Puting National Park, Borneo

Long-legged centipedes (Scutigera spp.) are ferocious predators of the forest floor – extremely swift footed with excellent vision and formidable needle-sharp fangs through which paralysing venom is injected to immobilise other forest arthropods and small vertebrates like frogs and lizards - they are unusual in having well developed, insect-like compound eyes (other centipedes have simple eyes). Danum Valley, Kalimantan, Borneo

A picture of light reflected and focused through dew drops suspended from a garden spider's, Araneus diadematus, web. Calne, Wiltshire, UK.

A picture of light reflected and focused through dew drops suspended from a garden spider's, Araneus diadematus, web. Calne, Wiltshire, UK.

Flower spikes of Yorkshire Fog, Holcus lanatus, create a purple haze in low summer sun. Calne, Wiltshire, UK.

A wild duck feather framed by circles of confusion. River Marden. Calne, Wiltshire, UK

Teasel, Dipsacus fullonum, framed by circles of confusion Marden River, Calne, Wiltshire, UK

A poppy field forms a blur of abstract colour as the camera pans across the scene. Calne Wiltshire, UK

Reeds reflected in the still waters of Loch Lomond, Scotland, UK

Lacewing, Chrysopa carnea, rests amongst petals of a white Chrysanthemum flower. Helensburgh, Scotland, UK.

A Preying Mantis peers from a large arrow root leaf Danum Valley, Kalimantan, Borneo

Like a stained-glass window, the wing of a mayfly lights up with spectacular iridescent colours. Calne Wiltshire, UK.

Black seed, red pod: Like a flag, the black velvet-coated seed of a rainforest tree, Sterculia coccinea, is contrasted against its bright red pod. Tanjung Puting National Park, Borneo

The spectacular iridescent colours produced by a thin film of oil spread over a water surface belie the polluting reality of the dangers of tourism in pristine environments. Loango National Park, Gabon

A Large Red Damselfly, Pyrrhosoma nymphula (Odonata, Coenagriidae). Peers from its perch on a grass stem. Calne, Wiltshire, UK.

A small jumping spider hides amongst the down of a dandelion clock. Algarve, Portugal.

Hunting at night, a Chevron Tarantula, Psalmopoeus cambridgei, feeds on a long-horned grasshopper or Katydid in Arima Valley, Trinidad. The Katydid's eyes are black in the daytime (light adapted) but appear red at night (dark adapted) as in this photograph.

A female net-casting spider, also known as the Ogre-faced spider, Dinopis subrufa, peers through the expandable web net she spins and holds in her front pair of legs ready to throw over prey like forest-floor cockroaches passing beneath. Rainforest Australia.

A queen Common Wasp, Vespula vulgaris, rests on Grape Hyacinth, Muscari neglectum, in spring while searching for suitable nest site usually within abandoned small mammal hole. Calne, Wiltshire, UK

The eyes of this burrowing Frog are located towards the top of its head so that it can look upwards. Danum Valley, Kalimantan, Borneo.

The eye of a Red-eyed Tree Frog, or red-eyed leaf frog - Agalychnis callidryas, with vertical light adapted pupil slit. Chiquibul Forest reserve. Belize

Having just metamorphosed from the water-breathing tadpole stage, froglets of the common frog, Rana temporaria, must breathe air and so they climb out of the pond onto any available aerial surface. Here, a chorus-line of froglets compete for space on a frond of floating pond weed. Calne, Wiltshire, England, UK

The common frog, Rana temporaria, is a conspicuous inhabitant of our school pond – I’d been photographing the pond's life throughout the year and this very small juvenile has just swallowed a worm nearly as big as itself. Pausing a moment, the frog waits for the worm to crawl itself further into its stomach so that it can seal its mouth shut to permanently trap the worm! The frog now appearing as though comfortably chewing on a fat cigar! Calne, Wiltshire, UK.

Inflated like a balloon, a Mexican Burrowing Toad, Rhinophrynus dorsalis, lets out a series of deep moan-like calls to attract a mate as rains end of the dry season in Chiquibul Forest Reserve, Belize.

A Red-eyed Tree Frog, or red-eyed leaf frog, Agalychnis callidryas, sits above a pond within a lily flower during the breeding season. Braulio Carrillio National Park, Costa Rica

Giant whip spider, Heterophrynus cheiracanthus (Chelicerata, Amblypygi). Despite their spectacular and fearsome appearance, whip spiders are not poisonous, and are dangerous only to their prey;- insects and other arthropods, frogs and small lizards, but not to humans. The long front-leg spines form a basket to trap prey against the spider's mouth. Central Ridge Forest Reserve, English Man’s Bay, Tobago, Trinidad & Tobago.

Dew drops suspended from the parachutes of a dandelion clock glisten like diamonds in the morning light. Calne, Wiltshire, UK

The fur of an endangered Gabon Black Colobus monkey, Colobus satanas anthacinus, smolders and shrouds the monkey in a blanket of acrid smoke during preparation for sale. Food Market, Libreville, Gabon.

The fur of an endangered Gabon Black Colobus monkey, Colobus satanas anthacinus, catches fire during preparation for sale. Food Market, Libreville, Gabon.

A man holds up an endangered Gabon Black Colobus monkey, Colobus satanas anthracinus, and an endangered West African Dwarf Crocodile, Osteolaemus tetraspis tetraspis, destined for the bush-meat trade. Food Market, Libreville, Gabon

The head of the male Stalk-eyed fly, Diopsis macrophthalma (Diptera, Diopsidae) is drawn out and elongated into long “stalks” on which sit his compound eyes. The idea here is that males fight each other for females at special Lekking or mating sites. Males face each other off and the male whose eyes stick out furthest (longest stalks) wins the fight and the female. Here the male is cleaning the surface of his right eye with special grooming hairs on his front legs. Loango National Park, Gabon.

Embodying the essence of "insectiness" this long-horned beetle (Coleoptera, Cerambycidae). peers over the edge of a rotten log where it has been laying its eggs. Queensland, Australia.

About 5 days after fertilization, a frog’s egg has developed into a complex wriggling, blind embryo with gills similar in form to those found in Humans, Birds, Fish and all vertebrates. Looking like an apostrophe, this is the tadpole of the Common Frog, Rana temporaria. Calne, Wiltshire, UK.

Brown Anole Lizard locally known as the Hu Wa, Anolis sagrei, on Thatch Palm, Thrinax morrisii. Thatch Palm is used to thatch buildings. Chiquibul Forest Reserve, Belize.

A mouse wanders through a frozen landscape formed by snow-covered strawberry plants in the dead of winter. St. Andrews, Scotland, UK.

Sleeping Eye: Seeing with your eyes closed - During the day, the eye of the Red-eyed tree frog or red-eyed leaf frog, Agalychnis callidryas, is closed, but the frog can still keep a look-out for danger through a web-like veil that masks the bright red eye from a predator’s sight. Chiquibul Forest Reserve, Belize.

A Chapman’s Zebra, Equus quagga chapmani, (a subspecies of the Plains Zebra, Equus quagga) shows his stripes. London Zoo, UK

A Cane Flower or Torch ginger flower, Etlingera sp., provides a valuable food source for birds and insects. Tropical rain forest of the highlands of Sao Tome. Sao Tome e Principe, West Africa.

The central female reproductive structures of the Magnolia, Magnolia stellata, flower curl in anticipation. Wiltshire, UK

A Painted Bronze-back tree-snake, Dendrelaphis pictus, flicks its red tongue to sense for danger. Danum Valley, Borneo.

An anole lizard, Anolis sp. (Iguanidae) peers through the clear panes of a glasswing butterfly, Greta oto. The transparent wings allow the butterfly to blend into its background and become invisible to potential predators. Chiquibul Forest Reserve, Belize. Central America

The toe pads of Gecko lizards (Reptilia, Squamata, Gekkonidae) are covered in millions of minute hairs which enable them to grip even the smoothest surface like glass, or even hang upside down from a ceiling. Here, an Australian species clings to glass revealing the dark bands of specialised hairs (not actually true "hair" as in mammals, but modified scales drawn into extremely fine hair-like structures) which grip the glass. Queensland Rainforest.

Capturing the essence of a stalking insect, a Preying Mantis prowls across the sunlit surface of a giant Arrow or Elephant Ear leaf. Main Ridge Forest Reserve, Tobago, Trinidad & Tobago.

A Queen buff-tailed Bumblebee, Bombus terrestris, hovers above a common field poppy, Papaver rhoeas, while packing collected poppy pollen into her pollen baskets attached to her hind legs. Calne, Wiltshire, UK

A garden snail, Helix dispersa, peers into the camera lens - extremely shallow depth of field has been used to throw the rest of the snail into a distant blur. Calne, Wiltshire, UK.

A Slow Worm - Anguis fragilis - (a legless lizard), blows a bubble of slug slime after swallowing its favourite food a grey garden slug. Calne, Wiltshire, UK.

An African swallowtail butterfly caterpillar, Papilio demodocus, flicks its red, smelly, forked Osmeterium in defence. Marantaceae forest, Mikongo, Lope National Park, Gabon

A Long-horned grasshopper, or Katydid, rests during the day hidden in the whorl of an Arrow-root (Marantaceae) plant. Marantaceae forest, Mikongo, Lope National Park, Gabon

When young, green Iguanas - Iguana iguana - are bright green in colour matching perfectly the tone of their surrounding vegetation and becoming invisible to potential predators. Central Ridge Forest Reserve, Tobago, Trinidad & Tobago.

The backlit wing feathers of the Tawny Owl, Strix aluco. Calne, Wiltshire, UK.

Number of Captions: 7

"Bush-meat" Wins 2nd place in 2007 Pictures of the Year International (POYi 65) General Division Science/Natural History http://www.poy.org/65/01/ index.php March 2008 "Bush-meat" wins place in PDN - Photography Annual 2007- 3 images February 2008 Two portfolios of 10 natural history photographs and 10 science photographs are short-listed (final 10 photographers) in NATURE and SCIENCE categories on the professional side of the SONY World Photography Awards 2007 October 2007 "Frog & Worm" wins a Best of Show Award in the International Photography Awards 2007 http://www.photoawards. com/bos.asp

October 2008 David is named "European Wildlife Photographer of the Year 2008" at GDT International Nature Photography Festival, Luenen, 25th Oct. 2008 http://www.gdtfoto.de/ Wildlife Photographer of the Year Awards - NHM London David will attend the awards ceremony and events for WPOY 28th & 29th Oct. & corporate opening of exhibition on 3rd Nov. 2008 WILDPHOTOS 08 David will be speaking at WildPhotos 08 Royal Geographical Society London, 31st Oct. & 1 Nov. www.wildphotos.org.uk

David is a plenary speaker in Kuusamo, Finland at the Kuusamo Nature Photo XIV 2009 from 11th - 13th September: www.kuusamo.fi/Resource.phx/sivut/sivut-kuusamo/ naturephoto/index.htx David joins the Wild Wonders of Europe dream team of nature photographers http://www.wild-wonders.com/ see interview: http://www.wild-wonders.com/the_photographers_featured.asp?show=47 Plantscapes Portfolio awarded “Best of Show” in IPA 2009 http://www.photoawards.com/en/Pages/Gallery/winner.php January 2009 National Geographic News Tuesday January 6th 2009 "Lizard-Snake Deadlock Steals the Show" "Deadlock" is the 2nd most viewed National Geographic photograph of 2008 See: http://news.nationalgeographic.com/news/2008/12/photogalleries/top-ten-photos/photo9.html

See David present “Still” at the Lunen Festival Saturday, October 29th 2011 Sunday October 30th 2011 http://www.gdtfoto.de/content.php?siteloc=42&action=open&owner=9 David is highlighted as one of the Top 50 Photographers as featured in PhotoPlus Magazine Issue 50 July 2011 http://www.myfavouritemagazines.co.uk/photography David joins the team to Judge the GDT Contest "European Wildlife Photographer of the Year 2011" http://www.gdtfoto.de/content.php?lang=en David Opens Solo Exhibition at the Hannu Hautala Nature Photography Center, Kuusamo, Finland : http://www.hannuhautala.fi/Resource.phx/sivut/sivut-hannuhautala/index.htx See David speak at the Scottish Nature Photo Fair 2009; Battleby, Perth 5th, 6th September: www.snh.org.uk/snpf/

October 2013 Microscopy Successes - David receives multiple awards at this years Nikon Small World, the Olympus Bioscapes, and the European Photographer of the Year (GDT) Awards. Join David at PhotoLive 2013 - 26-27 October, London www.photo-live.com to get tips and secrets to improve your Macro Photography See you all at the BBC/NHM Wildlife Photographer of the Year Awards Dinner 15th Oct.2013. Join David (with the author Sara Maitland) in a “Nature and Awe” presentation as part of the Southbank London Literature Festival 27th May 2013 at 12.00pm http://www.southbankcentre.co.uk/whatson/nature-awe-73670 Successes in the Nikon Small World Photomicrography Competition 2011 :- An Honorable Mention and two Images of Distinction http://www.nikonsmallworld.com/

October 2014 David will be at the 50th anniversary Wildlife Photographer of the Year Awards ceremony 21st Oct. Successes also in this year’s British Wildlife Photographer of the Year, Olympus Bioscapes, and the European Photographer of the Year (GDT) Awards. See David’s Print Exhibition “Imago” at the International Nature Photography Festival: Lunen Oct. 24th-26th. http://www.gdtfoto.de/seiten/exhibition-hall.html David is again the specialist photographer for the BBC documentary series with Prof. Brian Cox: This time for the highly acclaimed series: Human Universe: www.bbc.co.uk/humanuniverse

See "Imago" at Lunen, Germany: http://www.gdtfoto.de/seiten/exhibition-hall.html 24th - 26th Oct 2014

Number of News Headlines: 1

Specialist photographer: www.bbc.co.uk/humanuniverse